Holdings
Uniqueness of the Collection
The Collection is one of the largest and most comprehensive collections (more than
2,000,000 specimens) in Colorado and the Rocky Mountain Region. Much of the material has
been identified by specialists to the specific or generic level, making it very valuable
in terms of quality of material. Examples are:
On the Unique and More Important Parts of the Collection
1) Aphid collection. The monographic works (Aphididae of Colorado and Aphids of the Rocky
Mountain Region) by Gillette and Palmer are the basis of one of the largest aphid
collections in the world.
2) The Natural Resource Ecology Laboratory Soil Arthropod Collection. The 9,000+ slide
collection is derived from three primary sources: (1) the International Biological
Program's Pawnee Site Soil Arthropod Reference Collection, (2) National Research Ecology
Laboratory Soil Arthropod Research Collection, and (3) the Konza Prairie Research Natural
Area LTER Reference Collections and collections from the Jornada LTER. These collections
were assembled by Faculty Affiliate, Dr. Dave E. Walter, University of Queensland,,
Australia, a well-known acarologist. He has described numerous species of Mesostigmata and
other mites from the above collections. Additionally, paratypes and voucher specimens for
studies detailing the feeding habits of more than 100 species of predatory, mycophagous,
and omnivorous soil arthropods are included in the collections.
3) Homoptera Collection. The 60 Cornell drawers contain one of the more comprehensive
collections in the western United States. Revisions and species descriptions by C. P.
Gillette, C.F. Baker, and M.W. Nielson have resulted in excellent holdings of North
American species of Cicadoidea. B. C. Kondratieff has in preparation a publication on the
cicadas of Colorado. The majority of North American species of cicadas is represented.
4) Diptera Collection. This section includes 195 Cornell drawers and is very comprehensive
in regard to coverage of most North American families. During revisions or species
descriptions and faunal surveys of this section has been greatly enriched by adding
secondary types and voucher material: Ceratopogonidae (M.T. James), Bibionidae (M.T.
James, S. Fitzgerald, and E. Hardy), Cecidomyiidae (W. Brewer), Stratiomyiidae (M.T.
James), Syrphidae (E.A. Back), Asilidae (E.A. Back), Mydidae (S. Fitzgerald, B.
Kondratieff and J. L. Welch), Pyrgotidae (B. C. Kondratieff and S. Fitzgerald),
Tephritidae (B.F. Quisenberry) and Chloropidae and Cuterebridae (C.W.Sabrosky). The
Mosquito Collection is one of the largest in the Rocky Mountains, and served as a basis
for Harmston and Lawson's Mosquitoes of Colorado (1976, U.S. Dept. Health, Education,
Welfare. CDC-Atlanta, Georgia). Scientists from the Center for Disease Control in Fort
Collins have also added important representatives and voucher specimens of species from
Guadalcanal, Philippines and China.
5) The Hymenoptera Collection. This collection consisting of 270 Cornell drawers is among
the most comprehensive, especially of the Rocky Mountain Region. Through the efforts of
H.E. Evans, representation of the Vespoidea, Scolioidea, Sphecidae and Pompilidae have
national importance, representing the majority of North American species. Evans and his
students have added many rare species through their study of the taxonomy and behavior of
the above groups. There are many voucher specimens, especially in Nyssoninae and
Philanthinae (Sphecidae), accumulated by the research of Evans and his students. This
includes a complete collection of beewolves (Philanthus), the basis of Evans and O'Neill's
book The Natural History and Behavior of North American Beewolves (1988, Cornell
University Press, Ithaca, New York).
6) Orthoptera Collection. This collection has comprehensive coverage of the Great Plains
species, including large series of economically important taxa. There are many voucher
specimens and paratypes by L. Bruner, J.L. Capinera, M. Hebard, A.P. Morse and J.A. Rehn.
The basis for J. L. Capinera and T. S. Sechrist's Grasshopper (Acrididae) of Colorado was
from the Collection.
7) The Coleoptera Collection includes 225 Cornell drawers of beetles significant North
American species of Cicindelidae, which was the basis of M. Kippenhan's Tiger Beetles
(Coleoptera: Cicindelidae) of Colorado, 1994. Trans. Amer. Entomol. Soc. 120: 1-86.
Elateridae, Scarabaeidae, Tenebrionidae, Cerambycidae (60% of all North American species
and basis for upcoming Checklist of Cerambycidae of Colorado in prep. by D. Heffern),
Chrysomelidae and Curculionidae.
8) The Lepidoptera Collection, under the curatorial egis of Dr. Paul A. Opler, includes
500 Cornell drawers and an excellent regional representation of Ditrysia, especially
Gelechioidea, Sesioidea, Tortricoidea, Pyraloidea, Papilionoidea, Geometroidea,
Bombycoidea, Sphingoidea, and Noctuoidea (the family Noctuidae alone includes 134 Cornell
drawers). More than 80% of all North American species of butterflies are represented. Many
of the specimens have been identified by experts such as R. Brown, D. R. Davis, D. C.
Ferguson, R. W. Hodges, J. McDunnough, R. W. Poole, J. A. Powell, and P. A. Opler. Many
specimens served as voucher material for Opler's books such as Peterson's Field Guide to
the Butterflies of Eastern North America and Western North America and distributional
atlases of western North American moths and eastern butterflies.
9) The aquatic insects (Ephemeroptera, 50% of North American species, many reared adults
and associated with larva; Plecoptera (90% of all North American species; and Trichoptera,
50% of all North American species) are one the fastest growing sections of the Collection.
A comprehensive coverage of western and many eastern species is now available: this part
of the Collection served and will serve as the basis for numerous manuscripts and books
(J. V. Ward and B. C. Kondratieff. 1992. An Illustrated Guide to the Mountain Stream
Insects of Colorado, University of Colorado Press). The mayfly material served as the
basis for one of the most comprehensive species inventories reported from any state (W. P.
McCafferty, R. S. Durfee and B. C. Kondratieff. 1993. Colorado Mayflies (Ephemeroptera):
an Annotated Inventory, Southwest. Nat. 38: 252-274). Additionally, many rare Neotropical
species are included, especially from Mexico.
10) Other taxa that are well represented include Hemiptera and other Homoptera, including
a comprehensive holding of North American Cicadoidea (80% of all species).
11) The Collection served as a repository for many of the insect species collected from
the Central Plains Experiment Range, Pawnee National Grassland, and IBP program funded by
NSF (Kumar, R., R.J. Lavigne, J.E. Lloyd, and R.E. Pfadt. 1976. Insects of the Central
Plains Experiment Range, Pawnee National Grassland. Agric. Exper. Sta. Univ. Wyoming.
Science Monogr. 32)
14) Thirty primary and over 2,500 secondary types, representing 9 orders and 50 families
(Odonata, Plecoptera, Orthoptera, Homoptera, Hemiptera, Diptera, Lepidoptera, Coleoptera,
Hymenoptera) are deposited in the Collection.
15) Approximately 90 percent or more of all North American species are represented in the
following groups: Odonata, Plecoptera, Orthoptera (s.s), many Hemiptera families, many
Homoptera, many Neuropteran families, many coleopteran families, Mecoptera, many dipteran
families, many trichopteran families, many lepidopteran families and many hymenopteran
families.
16. The repository for comprehensive arthropod inventories of Canyonlands National Park,
Great Sand Dunes National Monument, Mesa Verde National Park, Rocky Mountain National
Park, Dinosaur National Monument, Colorado National Monument and Zion National Park. These
studies have resulted in three fully illustrated outreach publications which are for sale
at each Park: A Guide to the Common Insects and Other Arthropods of Colorado National
Monument, Insects and other Arthropods of Great Sand Dunes National Monument and Kingdom
of the Small, Common Insects and Other Arthropods of Dinosaur National Monument.
Approximately, 7,500 species of arthropods were deposited into the Collection.