| Title |
Investigators | Department | Objectives | Approach Keywords | Progress Reports | Impact Statements | Publications | |
Project * COL00795 | |
| Title | Improvement of Quality and Performance of Colorado Wheat |
| Investigator(s) | Haley, SD; Johnson, JJ; Peairs, FB; |
| Department | Soil and Crop Sciences |
| Objectives | 1) To make available, through breeding and testing, winter wheat cultivars and germplasm having desirable end-use quality characteristics; 2) To make available, through breeding and testing, winter wheat cultivars and germplasm having desirable agronomic characteristics, 3) To conduct basic and collaborative research studies focused on mitigating constraints for effective production and marketing of wheat produced in Colorado. |
| Approach | Obtain a wide diversity of germplasm from domestic and international sources. Determine adaptability to Colorado conditions and use in hybridization to develop improved winter wheat cultivars and germplasm with the following characteristics: high yield and yield stability, good end-use quality, (milling, bread-making, Asian noodles), disease and insect resistance, environmental stress tolerance, and optimum agronomic adaptation for both dryland and irrigated production conditions. Conduct basic research relative to the genetic control of traits important to minimize production constraints and optimize end-use quality . Develop and evaluate methods of disease and insect resistance evaluation and bread-making and noodle quality. |
| Keywords | wheat breeding, milling quality, baking quality, noodle quality, insect resistance, disease resistance, high temperature tolerance, drought stress tolerance, Russian wheat aphid resistance |
| Progress Reports | |
| 1993 | Two advanced HRWW lines, CO880210 and CO880169, performed very well in 1991, 1992, and 1993 Colorado tests, and along with two RWA-resistant lines, are under increase for possible release in 1994. Selection progress was made for grain yield, grain volume weight, winter hardiness, resistance to shattering, drought tolerance, WSMV resistance, and bread-making quality. Cultivar performance trials and Russian wheat aphid evaluations were conducted statewide. Field studies to determine the economic injury level on a resistant wheat line were conducted at Fort Collins in 1992 and 1993. RWA infestations resulted in significant yield reduction on susceptible wheat, but not on the improved resistant wheat. The 4R chromosome of PI 386148 triticale was backcrossed into wheat in an unstable condition including small segments and single doses, whereas in PI 386150 triticale x Lamar, the rye chromosome 4R was found as a disomic substitution and/or addition. Progeny derived from crosses between PI 386150 and Lamar and PI 386148 x Lamar were selected only for resistance to RWA. Preferential retention of 4R in resistant progenies indicates that this chromosome is associated with RWA resistance. Expression of this resistance appears to be disturbed by the action of the wheat chromosomes and the dosage effect. The lack of rye/wheat chromosome interchanges in advanced generations may indicate little homoeology between S. montanum and wheat chromosomes; hence stabilization of resistance will be difficult. |
| 1994 | Three new cultivars were released in 1994; Akron and Halt are HRW wheats, and Sylvan is a HRS wheat. Akron is a semidwarf with very lax spikes. Compared to current cultivars, it has higher yield, hail tolerance, leaf rust tolerance, and excellent quality. Halt is the first Russian wheat aphid (RWA) - resistant wheat developed in the USA. It is an early semidwarf with excellent quality, is moderately susceptible to leaf rust, and is slightly lower yielding than Yuma. Sylvan is a semidwarf developed in Utah and has an 11% yield advantage over other wheats when grown under irrigation in southwestern Colorado. The first uniform RWA field screening test confirmed greenhouse screening tests results, showed that resistance levels of many new lines equal that of the resistant parents, and that there is a significant association between grain yield and RWA resistance. Early generation among-cross selection for chemical desiccation tolerance and the associated use of mechanical mass selection were studied. A cross between tolerant parents showed lower desiccation injury than crosses between sensitive parents or among contrasting parents. Mechanical mass selection within populations advanced with two cycles of chemical desiccation produced F4-bulk populations with significantly lower dessication injury. |
| 1995 | Halt, the first Russian wheat aphid (RWA)-resistant wheat developed in the USA, outyielded the predominant 'TAM 107' in 1995 variety tests. It is an early semidwarf with excellent quality, and is moderately susceptible to leaf rust. Selection progress was made for grain yield, grain volume weight, winter hardiness, resistance to shattering, drought tolerance, WSMV resistance, and bread-making quality. Cultivar performance trials and Russian wheat aphid evaluations were conducted statewide. A laboratory leaf unrolling test was able to detect differences between RWA resistant and susceptible wheats, but needs further modification before it would be suitable for large-scale RWA screening. Inheritance and allelism studies using seven new sources of RWA resistance indicate that six lines have single gene resistance and one has resistance conditioned by two genes. The new germplasm lines from Kansas and Oklahoma have genes already available in other lines, but five new lines had genes different from Dn4, 5 and 6. |
| 1996 | Halt, the first Russian wheat aphid (RWA)-resistant wheat developed in the USA, yielded equal or superior to predominant cultivars in 1996 on-farm tests in Colorado, Wyoming and western Nebraska. It is an early semidwarf with excellent quality, and is moderately susceptible to leaf rust. Akron, our newest release, was the highest yielding cultivar in the 1996 Colorado dryland variety tests. Selection progress was made for grain yield, grain volume weight, winter hardiness, resistance to shattering, drought tolerance, WSMV resistance, and bread-making quality among about 2000 early generation lines evaluated statewide. Molecular markers have been associated with the Dn2 and Dn4 genes conditioning RWA resistance. These markers will help shorten the development time for pyramiding resistance genes into adapted wheats. Genetic analyses of RWA resistance in the Russian triticale PI386150 showed single dominant gene resistance, and that the gene for resistance is located on chromosome 4R from S. montanum. Expression of the resistance gene was disturbed in a wheat background and no wheat-rye chromosome interchanges were found in advanced generations; therefore, stabilization of resistance will be difficult without artificial induction of chromosome interchanges. |
| 1997 | The objective of this project is to develop new cultivars and characterize genetics of important wheat traits. Selection progress was made for grain yield, grain volume weight, winter hardiness, resistance to shattering, drought tolerance, WSMV resistance, and bread-making quality among about 2000 early generation lines evaluated statewide. Resistance to RWA was determined by field screening of 8000 lines using artificial infestation. Molecular markers have been associated with the Dn2 and Dn4 genes conditioning RWA resistance. These markers will help shorten the development time for pyramiding resistance genes in adapted wheats. Two new wheats Yumar and Prowers, were released in 1997. They are resistant to the Russian wheat aphid, and are the first of their kind to be released after CSU's 1994 unveiling of 'Halt' , the first wheat resistant to the Russian wheat aphid introduced to United States farmers. Yumar and Prowers, were developed over eight years by backcrossing the resistance from an old Soviet wheat into two popular CSU varieties Yuma and Lamar. Yumar and Prowers will be available to farmers by next fall and are similar to Yuma and Lamar, their parent varieties . The new varieties should quickly replace their parent varieties and greatly broaden the adaptation of resistant wheats in Colorado. The RWA-resistant line TAM 107-R3 is under seed multiplication for probable release in August 1998. It was the highest yielding entry in both the 1997 low moisture dryland and 1997 high moisture dryland trials. It's milling, mixing, and baking quality has been similar to TAM 107. It contains about 90 percent RWA-resistant plants and is being further purified. The cultivars released from the program have occupied significant acreages (more than 20% of the 1997 crop, and 3 of the 4 leading cultivars). Assuming that Halt will occupy at least 10% of the acreage for each of 5 years (total 1 .4 million acres), and the savings per acre for not using insecticide is $10, the total benefit of Halt will e $14 million. Cost of development of Halt was estimated at $500,000. With the release of Yumar and Prowers, the average annual loss of about $10 million due to RWA control should be eliminated, and the environment protected from dangerous insecticides. The spring wheat breeding program began in 1996, and the major objectives are RWA resistance, heat tolerance, and early maturity. The earliest cultivar release date is 2003. The F3's are currently in the greenhouse SSD program. A herbicide resistance program began in 1997 with a grant from American Cyanamid. The first crosses were made during July 25-28, 1997 and depending on the acceleration of the program, lines could be ready for release in August 2002, 2003 or 2004. |
| 1998 | The objective of this project is to develop new cultivars and characterize the genetics of important wheat traits. Selection progress was made for grain yield, grain volume weight, winter hardiness, resistance to shattering, drought tolerance, WSMV resistance, and bread-making quality among about 2000 early generation lines evaluated statewide. Resistance to RWA was determined for the breeding program by field screening of about 8000 lines using artificial infestation. Molecular markers have been associated with the Dn2 and Dn4 genes conditioning RWA resistance; however, they are not close enough to the gene of interest to be useful in breeding. A new wheat, Prairie Red, was released in 1998. It is resistant to the Russian wheat aphid, and very similar to TAM 107 in all other traits. Prairie Red was developed over nine years by backcrossing the resistance from an old Soviet wheat into a popular cultivar TAM 107. Prairie Red will be available to farmers by next fall. The new variety, in addition to Halt, Yumar and Prowers, should quickly replace their parent varieties and greatly broaden the adaptation of resistant wheats in Colorado. Significant breeding advances have been made and host plant resistance has become the key to integrated management of the RWA. An understanding of the mechanisms of resistance associated with the major resistance genes, and/or molecular markers associated with them, will be very valuable in developing durable resistance through gene pyramiding and deployment. Three molecular markers have been reported, and other studies are underway. The spring wheat breeding program began in 1996, and the major objectives are RWA resistance, heat tolerance, and early maturity. The earliest cultivar release date is 2003. The F6's are currently being screened for RWA resistance and heat tolerance and the survivors will be field planted in March 1999. A herbicide resistance program began in 1997 with a grant from American Cyanamid. The first crosses were made during July 25-28, 1997 and depending on the acceleration of the program, lines could be ready for release in August 2002, 2003 or 2004. Doubled haploids have been successfully used to produce pure lines which will be planted in Fall 1999. |
| 1999 | The overall objectives of this project were to develop wheat cultivars with improved end-use quality and field performance and to conduct basic research related to winter wheat genetic improvement and production. Five winter wheat cultivars have been developed and released since 1995: Yumar and Prowers (1997), Prairie Red (1998), and Prowers 99 and Hayden (1999). Aside from Hayden, an introduction from Idaho released for western-slope dryland production, these cultivars each possess the Dn4 resistance gene which provides excellent protection against the Russian wheat aphid (RWA) and eliminates the need for costly and environmentally-unsound insecticide treatments. Basic RWA resistance research has provided valuable genetic information for alternative RWA resistance genes, which will eventually be deployed (either alone or in combination with Dn4) to ensure durability of RWA resistance in Colorado. Basic research on high temperature and drought stress provided useful genetic information on traits predictive of tolerance, including 2,3,5-triphenyltetrazolium chloride (TTC) reduction, membrane thermal stability, and chemically-induced nonstructural carbohydrate remobilization. Adoption of these procedures as routine screening criteria in the breeding program will hopefully lead to improved high temperature and drought stress tolerance in both spring and winter wheat cultivars. A hard white winter wheat (HWW) breeding effort, initiated in 1994, focused on incorporation of RWA resistance into the HWW germplasm base and adopting procedures for noodle-quality evaluation. Cultivar releases are expected in 2001 and 2004, allowing Colorado producers access to the very rapidly expanding export market for Asian noodles. A spring wheat breeding program, initiated in 1996, is currently focused on improved high temperature tolerance, early maturity, and RWA resistance. Advanced breeding lines will be planted in spring 2000 for the earliest cultivar release date in 2003. A wheat-maize dihaploid production system was adopted in 1997 as a complement to the overall breeding effort. Using this system, lines carrying RWA resistance and proprietary, non-transgenic tolerance to imidazilonone herbicides were developed and are positioned for cultivar release in 2002. Tolerance to imidazilonone herbicides would allow for selective control of winter annual grassy weeds (e.g., Jointed goatgrass, downy grome, cheatgrass) that are responsible for $25 million annual yield losses in Colorado. |
| 2000 | The primary objective of this project is to develop wheat cultivars and germplasm having desirable agronomic and end-use quality characteristics. Three advanced lines performed very well in 2000 and are under increase for possible release in 2001. Two of these lines (CO980889 and CO980894) carry non-transgenic tolerance to imidazolinone herbicides which would allow selective control of winter annual grassy weeds (e.g., jointed goatgrass, cheatgrass, downy brome) that are problematic in Colorado and other wheat production areas. The third line under large-scale increase (CO940611) is a hard white winter wheat (HWW) experimental line with high yield and superior milling and bread making characteristics. Eight elite breeding lines were advanced for breeder seed increase in 2001 to enable potential foundation seed increase in 2002. These lines include: five lines with Russian wheat aphid (RWA) resistance in an 'Akron' background; one line (CO950043) with superior dryland and irrigated yield and a previously undeployed RWA resistance gene; one line (CO970498) with RWA resistance and superior yield and milling and baking quality; and one line (CO970547) with high yield, RWA resistance, and barley yellow dwarf virus (BYDV) tolerance. Three lines derived from a wheat x maize doubled haploid program, carrying both non-transgenic herbicide tolerance and RWA resistance, were advanced for testing in state variety trials in 2001. Significant progress was realized in development and adoption of small-scale tests for prediction of key noodle quality characteristics in HWW, and several experimental lines with RWA resistance and good dual purpose (bread and Asian noodles) were advanced from preliminary yield trials to the replicated yield testing stage. To provide basic information and tools to enhance the breeding efforts, the following research projects were initiated or continued in 2000: inheritance and chromosomal location of a novel wheat streak mosaic virus (WSMV) gene; refinement and improvement of wheat x maize doubled haploid development procedures; characterization of advantages and disadvantages of incorporation of a Gibberellic acid-sensitive semidwarfing gene (Rht8); determination of genotypic and environmental influences on hard white wheat quality characteristics; and stem carbohydrate remobilization in relation to postanthesis stress tolerance. Spring wheat advanced lines were tested at 3 sites in eastern Colorado; however, severe drought destroyed all trials. |
| 2001 | The objectives of this project are to a) develop wheat cultivars and germplasm having desirable agronomic and end-use quality characteristics and b) conduct basic research to improve understanding of genetic and environmental factors that affect wheat yield and end-use quality. Three advanced lines on foundation seed increase were released to seed producers as improved cultivars in fall 2001. Two of these, named ABOVE and AP502 CL, are hard red winter wheat (HRW) cultivars that carry non-transgenic tolerance to the new imidazolinone herbicide BEYOND from BASF Corporation. The first publicly developed winter wheat cultivars of their kind, ABOVE and AP502 CL will allow selective control of winter annual grasses (e.g., goatgrass, brome and cheat, and feral rye) and broadleaf weeds that are problematic in Colorado and other wheat production areas. The third cultivar released to seed producers, named AVALANCHE, is a hard white winter wheat (HWW) with high yield and superior milling and bread making characteristics. Adoption and production of HWW cultivars such as AVALANCHE in Colorado and the west central Great Plains may provide wheat producers with additional marketing options to increase farm profitability. Two lines that carry Russian wheat aphid (RWA) resistance backcrossed into the widely popular winter wheat cultivar AKRON were advanced for further field testing; foundation seed increase is being done to enable potential release of one line as an improved cultivar in Fall 2002. Spring wheat cultivar development, initiated in 1996 to provide growers with additional spring cropping options for reduced tillage production systems, progressed to identification of 12 experimental lines combining good agronomic characteristics (e.g., earliness, high temperature tolerance) with RWA resistance. These lines will be evaluated in replicated field trials at multiple locations in 2002 with simultaneous preliminary seed increase to enable potential release of an improved cultivar in spring 2004. Significant progress was realized in development and adoption of small-scale tests for prediction of key noodle quality characteristics in HWW and experimental lines with RWA resistance and good dual purpose (bread and Asian noodles) were advanced from advanced yield trials to the replicated yield testing in state variety trials in 2002. Continued development of adapted herbicide tolerant wheat cultivars also progressed to advancement of lines combining herbicide tolerance, RWA resistance, and improved milling and baking quality for yield testing in state variety trials in 2002. To provide basic information and tools to enhance the breeding efforts, the following collaborative research projects were continued or initiated in 2001: inheritance and chromosomal location of a novel wheat streak mosaic virus (WSMV) resistance gene; determination of breeding potential of gibberellic acid-sensitive semidwarfing genes; determination of genotypic and environmental influences on hard white wheat quality characteristics; characterization of antioxidant properties of wheat grain; inheritance and allelism of RWA-resistance in Iranian landrace accessions. |
| 2002 | The objectives of this project are to a) develop wheat cultivars and germplasm having desirable agronomic and end-use quality characteristics and b) conduct basic research to improve understanding of genetic and environmental factors that affect wheat yield and end-use quality. In Fall 2002, one advanced line was released as a new cultivar named 'Ankor'. Ankor was derived by backcrossing Russian wheat aphid (RWA) resistance into the popular winter wheat cultivar 'Akron'. Field and laboratory tests have shown that Ankor has similar dryland yield, higher irrigated yield, higher kernel weight and more uniform kernel size distribution, and better baking quality as compared to Akron. Development of cultivars such as Ankor provides wheat producers in Colorado and the west-central Great Plains with an effective, economical, and environmentally-sound means of mitigating economic losses from RWA. Spring wheat cultivar development, initiated in 1996 to provide additional cropping options for reduced tillage production systems, continued with the testing of 12 advanced lines combining earliness, high temperature tolerance, and RWA resistance at four Colorado locations. Due to severe drought and high temperature conditions, no data were obtained from these trials. Seed increases grown under irrigation will allow evaluation of these lines in 2003 to enable potential release of an improved cultivar in 2005 or 2006. Through a federally-funded grant project (USDA-IFAFS), two cycles of backcross marker-assisted selection were completed to transfer genes for viral resistance, RWA resistance, and high grain protein content to elite breeding lines and cultivars. Adoption of these molecular-DNA markers will foster the rapid development of wheat cultivars with new pest resistance traits and enable pyramiding of multiple RWA resistance genes into the same cultivar to provide long-term stability of resistance. Experimental hard white wheat lines with RWA resistance and good dual purpose quality (bread and Asian noodles) were selected for yield testing in state variety trials in 2003. Development of herbicide tolerant cultivars also progressed to selection of lines combining herbicide tolerance, RWA resistance, and improved milling and baking quality for yield testing in variety trials in 2003. Master's thesis research on a novel wheat streak mosaic virus resistance gene suggested inheritance by one dominant and one recessive gene; data on chromosomal location(s) of the resistance were inconclusive. To provide basic information to enhance the breeding efforts, several collaborative research projects were continued or initiated in 2002: environment and genotype x environment interaction effects on Asian noodle quality; breeding potential of gibberellic acid sensitive semidwarfing genes that do not reduce coleoptile length; geographic information systems (GIS) technology to improve variety recommendations and identify variety-specific production and quality zones; separation of the Dn7 RWA resistance gene from deleterious wheat quality factors; characterization of RWA resistance in Iranian landrace selections; characterization of antioxidant properties of wheat grain. |
| 2003 | The objectives of this project are to a) develop wheat cultivars and germplasm having desirable agronomic and end-use quality characteristics and b) conduct basic research to improve understanding of genetic and environmental factors that affect wheat yield and end-use quality in Colorado. In fall 2003, two experimental lines (CO980607 and CO00D007) were advanced for Foundation seed production to enable release as new cultivars in fall 2004. CO980607 (Yuma/T-57//TAM 200/3/4*Yuma/4/KS91H184/Vista pedigree) is a medium maturity, semidwarf, hard red winter wheat with good test weight, good shattering tolerance, moderate resistance to stripe rust, resistance to Russian wheat aphid (biotype A), and excellent overall milling and baking properties. CO980607 was the highest yielding entry in the 2003 Colorado Uniform Variety Performance Trials (UVPT) and is second only to the cultivar Trego on a three-year average. CO00D007 (Yumar//TXGH12588-120*4/FS2 pedigree) is non-transgenic herbicide tolerant ("Clearfield" wheat), medium-early maturity, tall semidwarf, hard red winter wheat with good shattering tolerance, resistance to Russian wheat aphid (biotype A), and excellent overall milling and baking properties. CO00D007 was the second highest yielding entry in the 2003 UVPT and is the highest yielding entry on a two-year average. In spring 2003 , a new biotype of RWA was identified in southeast Colorado. This new biotype (provisionally denoted as "biotype B"), the first reported variant in North American RWA populations, is virulent on all RWA-resistant cultivars available in the US. A search for germplasm sources resistant to the new biotype is underway and several promising sources have been identified for immediate use in breeding resistant cultivars. Through a federally-funded grant project (USDA-IFAFS), one cycle of backcross marker-assisted selection was completed to transfer genes for viral resistance, RWA resistance (Dn2 and Dn4 resistance genes), and high grain protein content gene to elite breeding lines and cultivars. The Dn2 and Dn4 backcross programs have been discontinued due to the appearance of the new RWA biotype. A federally-funded grant project (USDA-CSREES Special Grant) was initiated to develop new technologies for cultivar development, specifically the identification of molecular markers linked with new RWA resistance sources, transfer of drought and high temperature tolerance factors from wheat relatives, and implementation of whole-grain near infrared spectroscopy (NIRs) for wheat quality assessment. To provide basic information to enhance the breeding efforts, several collaborative research projects were continued or initiated in 2003: breeding potential of gibberellic acid sensitive semidwarfing genes that do not reduce coleoptile length; geographic information systems (GIS) technology to improve variety recommendations and identify variety-specific production and quality zones in Colorado; separation of the Dn7 RWA resistance gene from deleterious wheat quality factors; characterization of RWA resistance in Iranian landrace selections; identification of wheat germplasm resources with resistance to the new biotype of RWA. |
| 2004 | Two new hard red winter wheat cultivars were released in fall 2004. This first of these, named 'Hatcher', combines high dryland yields in eastern Colorado, resistance to Biotype 1 Russian wheat aphid (RWA), and improved milling and baking quality relative to other available RWA-resistant cultivars. Hatcher was selected from the crosses and backcrosses Yuma/PI 372129//TAM-200/3/4*Yuma/4/KS91H184/Vista. Across 22 trial locations of the dryland Colorado UVPT (2001-2004), Hatcher had lower yield than Trego but higher yield than all other entries in the trials. Hatcher has shown excellent milling properties and above-average bread baking properties. The second of the new cultivars, named 'Bond CL', combines resistance to Biotype 1 RWA, the Clearfield herbicide tolerance gene, and excellent bread baking quality. Bond CL is a doubled-haploid line developed using the wheat x maize hybridization method from the cross Yumar//TXGH12588-120*4/FS2. Across 14 trial locations of the dryland Colorado UVPT (2002-2004) , Bond CL had lower yield than Above Clearfield wheat and Hatcher but higher yield than all other entries in the trials. Bond CL has shown much-improved baking quality characteristics relative to Above. We continued our search for germplasm with resistance to the new biotype of RWA identified in 2003 (designated as 'Biotype 2'). We confirmed biotype 2 resistance identified in 2003 and have made crosses with these materials for both breeding purposes and mapping population development. We also screened over 5,000 Iranian landrace selections and have identified many accessions with biotype 2 resistance as effective as that expressed in the 94M370 germplasm line (Dn7 gene). In spring 2004, additional virulent RWA biotypes were identified in Colorado and wheat production areas in adjacent states. Working in collaboration with scientists in Oklahoma, we have confirmed that some of the most promising biotype 2 resistance sources (except the Dn7 gene) also confer resistance to one or more of the new biotypes. Field research confirmed that the Dn7 gene provided as much protection from RWA as the Dn4 gene under artificial RWA biotype 1 infestation. A thesis project was completed focusing on geographic information systems (GIS) technology to improve variety recommendations and identify variety-specific production and quality zones in Colorado. Wheat production areas of eastern Colorado were split in two mega-environments: a high precipitation, high yielding zone in the northeast part of the state and a low yielding zone characterized either by low precipitation along the Front Range or by high temperatures in the Southeast. Significant genotype x environment interaction was revealed between the two mega-environments, which is useful to facilitate interpretation of variety response in Colorado and refining variety recommendations. |
| 2005 | In fall 2005, one experimental line (CO00016) was advanced for Foundation seed production to enable release in fall 2006. CO00016 was developed from the cross CO940606/TAM107R-2 made in 1996. CO00016 is a white-chaffed, short semidwarf (similar height and heading date as Prairie Red) with excellent high temperature and drought stress tolerance. CO00016 is resistant to biotype 1 of the Russian wheat aphid (RWA), susceptible to biotype 2 of RWA, susceptible to both leaf and stripe rust, and moderately susceptible to stem rust. In Colorado dryland variety trials, CO00016 has shown high yields, excellent yield stability, and average test weights (similar to Jagger and Yuma). CO00016 is currently in its fourth year of statewide testing. Across 21 dryland trial locations between 2003 and 2005, CO00016 has been the highest yielding entry in the trials, about 0.5 bu/a higher than Bond CL, 1.7 bu/a higher than Hatcher, 3.1 bu/a higher than Above, 4.1 bu/a higher than Avalanche, and 5.3 bu/a higher than Jagalene (the next five highest yielding entries following CO00016). CO00016 has also shown very good milling and bread baking quality characteristics based on testing in the CSU Wheat Quality Lab and the USDA-ARS Hard Winter Wheat Quality Lab (Manhattan, KS). In the breeding program, we continue to focus on several different research areas. Resistance to new Russian wheat aphid (RWA) biotypes continues to be used in the crossing program, both for forward (three-way crosses) as well as backcrossing into elite backgrounds. We completed the screening of 7,300 Iranian landrace selections for resistance to RWA biotype 2 with approximately 330 biotype 2 resistant accessions also showing resistance to RWA biotype 1. Approximately 155 accessions carry resistance to both biotypes; DNA molecular marker mapping populations are under development with at least 5 of these accessions. Three advanced experimental lines with resistance to RWA biotype 2 derived from Triticale performed well enough to justify advance to state variety trials in fall 2005. Cytological testing of these lines, performed by Dr. Kabwe Nkongolo in Canada, suggest that the chromosomal regions responsible for the resistance are stable. Clearfield wheat breeding efforts involved advance of several promising hard white wheat experimental lines to state variety trials and rapid incorporation of a second gene for tolerance into our germplasm base. Stripe rust again was a significant production concern in 2005. A very high level of infection was observed at Fort Collins and resistance ratings from breeding materials figured prominently in selection decisions. A dissertation project was completed focusing on characterization of allelic diversity and associated agronomic effects of variation at the Xgwm261 microsatellite marker locus in wheat. |
| 2006 | In fall 2006, experimental line CO00016 was released as an improved cultivar named Ripper. Ripper was developed from the cross CO940606/TAM107R-2 made in 1996. Ripper is a white-chaffed, short semidwarf (similar height and heading date as Prairie Red) hard red winter wheat. Ripper has shown high dryland yields, excellent milling and bread baking quality, and excellent drought and high temperature tolerance. In four years of statewide testing in the dryland Colorado Uniform Variety Performance Trial (UVPT), Ripper was the top yielding entry in the trials, approximately 1.6 bu/a greater than Hatcher, 3.8 bu/a greater than Yuma, and 4.2 bu/a greater than Prairie Red. Ripper is resistant to biotype 1 of the Russian wheat aphid (RWA), susceptible to biotype 2 of RWA, susceptible to both leaf and stripe rust, and moderately susceptible to stem rust. Ripper will be an excellent replacement for other stress tolerant, early-maturing cultivars in Colorado, particularly TAM 107 and Prairie Red, which tend to perform better in dry years yet have a marketing penalty due to their poor milling and bread baking quality. In fall 2006, one experimental line (CO01385-A1) was advanced for Foundation seed production to enable release in fall 2007. CO01385-A1 was developed from the cross Yumar/Arlin made in 1997. CO01385-A1 is a white-chaffed, medium-height semidwarf with high yield under both dryland and irrigated conditions, high test weight, and good stripe rust resistance. CO01385-A1 is currently in its third year of statewide testing. Across 21 dryland trial locations between 2005-2006, CO01385-A1 has been the highest yielding entry in the trials, 0.8 bu/a greater than Ripper, 1.7 bu/a greater than Bond CL, 2.9 bu/a greater than Hatcher, 3.7 bu/a greater than Keota, and 4.4 bu/a greater than Avalanche (the next five highest yielding lines). Test weight of CO01385-A1 in these trials was the third-highest among all entries, about 1 lb/bu greater than the trial average. CO01385-A1 was the highest yielding entry averaged across two years (six total locations) of testing in Irrigated Variety Performance Trial, about 3.1 bu/a greater than TAM 111, 4.9 bu greater than Bond CL, and 9.2 bu greater than Hatcher (the next three highest yielding lines) . CO01385-A1 has shown lower grain protein and average bread baking quality charistics based on testing in the CSU Wheat Quality Lab and the USDA-ARS Hard Winter Wheat Quality Lab (Manhattan, KS). Resistance to new RWA biotypes continues to be used extensively in the crossing program, both for forward (three-way crosses) as well as backcrossing into elite backgrounds. Molecular marker mapping populations have been developed with several of these sources and mapping is currently underway with two different sources. An advanced trial of experimental RWA biotype 2 resistant lines was planted in fall 2006 at several Colorado locations; entries were included from our program and the Oklahoma State University breeding program. Clearfield wheat breeding efforts involved advance of a group lines carrying both tolerance genes to unreplicated field trials at our main testing sites. |
| Impact | |
| 1999 | The development and release of RWA-resistant cultivars will essentially eliminate yield and quality losses, as well as pesticide expenses and their environmental effects when they are used to control RWA. |
| 2000 | The primary objective of this research is to develop and release improved wheat cultivars and germplasm with enhanced grain yield, multiple pest resistance traits, and desirable end-use quality characteristics. Development of improved wheat cultivars will help wheat producers and the wheat industry in Colorado and the west central Great Plains by reducing costs of wheat production, minimizing the use of chemical pesticides, and allowing more successful marketing of wheat. |
| 2001 | The primary objective of this research is to develop and release improved wheat cultivars and germplasm with enhanced grain yield, multiple pest resistance traits, and desirable end-use quality characteristics. Development of improved wheat cultivars serves a vital function for wheat producers and the wheat industry in Colorado and the west central Great Plains by reducing costs of wheat production, minimizing the use of chemical pesticides, and allowing more successful marketing of wheat to improve farm profitability. |
| 2002 | The primary objective of this research is to develop and release improved wheat cultivars and germplasm with enhanced grain yield, multiple pest resistance traits, and desirable end-use quality characteristics. Development of improved wheat cultivars serves a vital function for the wheat industry in Colorado by reducing costs of wheat production, minimizing or eliminating the need for chemical pesticides, and providing improved wheat marketing options. Since inception of the program in 1963, the CSU Wheat Breeding Program has released more than 20 improved wheat cultivars. CSU-bred wheat cultivars account for nearly 60% of Colorado's 2.6 million acres (2003 crop) with the remaining acreage planted almost exclusively with cultivars from university breeding programs in adjacent states. Estimates from Colorado wheat industry leaders indicate that quality enhancements from these wheat cultivars provide an average of $20 million per year increased income for Colorado wheat producers (83 million bushels x $0.25 per bushel price increase; 2003 dollars). CSU-bred Russian wheat aphid resistant cultivars are currently planted on over 25% of the Colorado acreage (2003 crop). Farmers who plant these cultivars and experience RWA infestations would save approximately $14 per acre in insecticide costs (one application only) and $22 per acre in reduced yield (average 25% yield reduction), a total benefit of over $23 million per year. |
| 2003 | The primary objective of this research is to develop and release improved wheat cultivars and germplasm with enhanced grain yield, multiple pest resistance traits, and desirable end-use quality characteristics. Development of improved wheat cultivars serves a vital function for the wheat industry in Colorado by reducing costs of wheat production, minimizing or eliminating the need for chemical pesticides, and providing improved wheat marketing options. Since inception of the program in 1963, the CSU Wheat Breeding Program has released more than 20 improved wheat cultivars. CSU-bred wheat cultivars account for nearly 60% of Colorado's 2.6 million acres (2003 crop) with the remaining acreage planted almost exclusively with cultivars from university breeding programs in adjacent states. Estimates from Colorado wheat industry leaders indicate that quality enhancements from these wheat cultivars provide an average of $20 million per year increased income for Colorado wheat producers (83 million bushels x $0.25 per bushel price increase; 2003 dollars). CSU-bred Russian wheat aphid resistant cultivars are currently planted on over 25% of the Colorado acreage (2003 crop). Farmers who plant these cultivars and experience RWA infestations would save approximately $14 per acre in insecticide costs (one application only) and $22 per acre in reduced yield (average 25% yield reduction), a total benefit of over $23 million per year. |
| 2004 | The primary objective of this research is to develop improved wheat cultivars with enhanced grain yield, multiple pest resistance traits, and desirable end-use quality characteristics. Development of improved wheat cultivars serves the wheat industry in Colorado by reducing wheat production costs, reducing pesticide use, and providing improved marketing options. Since inception of the program in 1963, the CSU Wheat Breeding Program has released nearly 25 improved wheat cultivars. CSU-bred wheat cultivars account for nearly 60% of Colorado's 2.4 million acres (2004 crop) with the remaining acreage planted almost exclusively with cultivars from university breeding programs in adjacent states. Estimates from Colorado wheat industry leaders indicate that quality enhancements from these cultivars provide an average of $20 million per year increased income for Colorado wheat producers (83 million bushels x $0.25 per bushel price increase; 2003 dollars). Russian wheat aphid resistant cultivars developed at CSU are currently planted on over 25% of the Colorado acreage (2004 crop). Farmers who plant these cultivars and experience RWA biotype 1 infestations would save approximately $14 per acre in insecticide costs and $22 per acre in reduced yield (average 25% yield reduction), a total benefit of over $23 million per year. Incorporation of resistance to new Russian wheat aphid biotypes identified in Colorado in 2003 and 2004 will provide wheat producers in affected areas with the protection needed to reduce pesticide use and achieve high yields in the presence of RWA. |
| 2005 | The objectives of the wheat breeding project are to a) develop wheat cultivars and germplasm having desirable agronomic and end-use quality characteristics and b) conduct basic research to improve understanding of genetic and environmental factors that affect wheat yield and end-use quality in Colorado. Development of improved wheat cultivars serves the wheat industry in Colorado by reducing wheat production costs, reducing pesticide use, and providing improved marketing options. Since inception of the program in 1963, the CSU Wheat Breeding Program has released over 25 improved wheat cultivars. CSU-bred wheat cultivars account for nearly 60% of Colorado's 2.4 million acres (2005 crop) with the remaining acreage planted almost exclusively with cultivars from university breeding programs in adjacent states. During this time, average wheat grain yields in Colorado have more than doubled with at least 50% of this increase attributed to improved cultivars. Estimates from Colorado wheat industry leaders indicate that end-use quality enhancements from cultivars developed at CSU provide an average of $20 million per year increased income for Colorado wheat producers (83 million bushels x $0.25 per bushel price increase; 2003 dollars). Production risks have been reduced significantly and the breeding program continues to address new production risks, such as those arising from the appearance of new Russian wheat aphid biotypes or races of stripe rust. |
| 2006 | The objectives of the wheat breeding project are to a) develop wheat cultivars and germplasm having desirable agronomic, disease and insect resistance, and end-use quality characteristics and b) conduct basic research to improve understanding of genetic and environmental factors that affect wheat yield and end-use quality in Colorado. Development of improved wheat cultivars serves the wheat industry in Colorado by reducing wheat production costs, reducing pesticide use, and providing improved marketing options. Since inception of the program in 1963, the CSU Wheat Breeding Program has released over 26 improved wheat cultivars. CSU-bred wheat cultivars account for over 50% of Colorado's 2.4 million acres (2006 crop) with the remaining acreage planted mostly with cultivars from university breeding programs in adjacent states. During this time, average wheat grain yields in Colorado have more than doubled with at least 50% of this increase attributed to improved cultivars. Estimates from Colorado wheat industry leaders indicate that end-use quality enhancements from cultivars developed at CSU provide an average of $20 million per year increased income for Colorado wheat producers (83 million bushels x $0.25 per bushel price increase; 2003 dollars). Production risks have been reduced significantly and the breeding program continues to address new production risks, such as those arising from the appearance of new Russian wheat aphid biotypes or races of stripe rust. |
| Publications | |
| 1993 |
HALEY, S.D., QUICK, J.S., AND MORGAN, J.A. 1993. Field excised- leaf water status evaluation and asscociations in winter wheat. Canadian J. Plant Sci. 73:55-63. KISANA, N.S., NKONGOLO, K.K., QUICK, J.S., AND JOHNSON, D.L. 1993. Production of doubled haploids by anther culture and the wheat x maize method in a wheat breeding program. Plant Breeding 110:96-102. KROENING, M.K., PEAIRS, F.B., QUICK, J.S., AND SHANAHAN, J.F. 1992. Economic injury level for Russian wheat aphid on a resistant wheat line in Colorado. Entom. Soc. Am. Annual Meeting. MORGAN, J.A., LECAIN, D.R., MCCAIG, T.N., AND QUICK, J.S. 1993. Gas exchange, carbon isotope discrimination, and productivity in winter wheat. Crop Sci. 33:178-186. NKONGOLO, K.K., QUICK, J.S., MUHLMANN, AND LAPITAN, N.L. 1993. An optimized fluorescence in situ hybridization procedure detecting rye chromatin in wheat. Genome 36:701-705. PEAIRS, F.B., QUICK, J.S., AND ECHOLS, J.E. 1992. Research progress on wheat aphid control. Colorado Wheat Farmer. 34(No.1):4. QUICK, J.S. 1993. Annual Wheat Newsletter. Vol. 39. 427 p. QUICK, J.S., SOUZA, E., AND SUNDERMAN, D.W. 1993. Registration of Fairview wheat. Crop Sci. 33:878. QUICK,J.S., NKONGOLO, K.K., AND PEAIRS, F.B. 1992. Breeding wheat for Russian wheat aphid resistance in the west central Great Plains. IN: Proc. 19th Hard Red Winter Winter Wheat Workers Conf., Lincoln, NE. |
| 1994 |
DONG, H. AND QUICK, J. S. 1994. Inheritance and Allelism of Russian wheat aphid resistance in a group of wheat lines. Agron. Abstracts, American Society of Agronomy, Madison, WI. p. 109 DONG, H., NKONGOLO, N. AND QUICK, J. S. 1994. Progress and problems in the transfer of Russian wheat aphid resistance from Russian triticales to wheat. Proc. Sixth Russian Wheat Aphid Workshop, Fort Collins, CO HALEY, S. D. AND QUICK, J. S. 1993. Early generation selection for chemical desiccation tolerance in winter wheat. Crop Sci. 33:1217-1223 KROENING, M., ET AL. 1994. Economic impact of Russian wheat aphid on resistant and susceptible wheats in Colorado. Proc. Sixth Russian Wheat Aphid Workshop, Fort Collins, CO QUICK, J. S. 1994. Annual Wheat Newsletter. Vol. 40. 380 p QUICK, J. S. 1994. Development of cultivars resistant to the Russian wheat aphid. Proc. Sixth Russian Wheat Aphid Workshop, Fort Collins, CO QUICK, J. S. AND DONG, H. 1994. First uniform Russian wheat aphid field test. Proc. Sixth Russian Wheat Aphid Workshop, Fort Collins, CO RANDOLF, T. ET AL. 1994. Tritrophic relationships: a comparison of TAM 107 and CORWA1 winter wheat varieties with Russian wheat aphid and Hippodamia variegata. Proc. 6th Russian Wheat Aphid Workshop, Fort Collins, CO SAIDI, A. AND QUICK, J. S. 1994. Inheriance of Russian wheat aphid resistance in 4 winter wheats. Proc. 6th Russian Wheat Ahid Workshop, Fort Collins, CO |
| 1995 |
DONG, H., AND QUICK, J.S. 1995. Detection of a 2.6 kb single/low copy DNA sequence on chromosomes of wheat (Triticum aestivum) and rye (Secale cereale) by flourescence in situ hybridization. Genome 38:246-249 DONG, H., AND QUICK, J.S. 1995. Inheritance and allelism of resistance to the Russian wheat aphid in seven wheat lines. Euphytica 81:299-303 DONG, H., QUICK, J.S., BRIGHAM, D.L., BJOSTAD, L.B., RUDOLPH, J.B., AND PEAIRS, F.B. 1994. Leaf unrolling of three wheat genotypes in Russian wheat aphid extracts. Cereal Res. Commun. 22:375-379 MORGAN, J.A., ZERBI, G., MARTIN, M., MUJAHID, M.Y., AND QUICK, J.S. 1993. Carbon isotope discrimination and productivity in winter wheat. J. Agronomy and Crop Science 171:289-297. |
| 1996 |
MA, Z.Q., NELSON, C., SAIDI, A., QUICK, J.S., AND LAPITAN, N.L.V. 1996. Chromosome localization and mapping of Russian wheat aphid resistance genes Dn2 and Dn4 in wheat. Genome IV. Scherago International, Inc., San Diego, CA. Abs. p. 149 NKONGOLO, K.K., LAPITAN, N.L.V., AND QUICK, J.S. 1996. Genetic and cytogenetic analyses of Russian wheat aphid resistance in triticale x wheat hybrids and progenies. Crop Sci. 36:1114-1119 QUICK, J.S., ALBRECHTSEN, R.S., AND STACK, M. 1996. Registration of Sylvan wheat. Crop Sci. 36:802 QUICK, J.S., AND REYNOLDS, M. 1996. Comparison of methods for selection for heat tolerance in wheat. Proc. 5th International Wheat Conference, Ankara, Turkey QUICK, J.S., ELLIS, G.E., NORMANN, R.M., STROMBERGER, J.A., SHANAHAN, J.F., PEAIRS, F.B., RUDOLPH, J.B., AND LORENZ, K. 1996. Registration of Russian wheat aphid-resistant wheat germplasm. Crop Sci. 36:217 QUICK, J.S., ELLIS, G.E., NORMANN, R.M., STROMBERGER, J.A., SHANAHAN, J.F., AND LORENZ, K. 1996. Registration of Akron wheat. Crop Sci. 36:210-211 QUICK, J.S., ELLIS, G.E., NORMANN, R.M., STROMBERGER, J.A., SHANAHAN, J.F., PEAIRS, F.B., RUDOLPH, J.B., AND LORENZ, K. 1996. Registration of Halt wheat. Crop Sci. 36:210 QUICK, J.S., ELLIS, G.E., NORMANN, R.M., STROMBERGER, J.A., SHANAHAN, J.F., AND LORENZ, K. 1996. Registration of Jules wheat. Crop Sci. 36:208 SAIDI, A., AND QUICK, J.S. 1996. Inheritance and allelic relationships among Russian wheat aphid resistance genes in winter wheat. Crop Sci. 36:256-258 |
| 1997 |
DONG, H., QUICK, J.S., AND ZHANG, Y. 1997. Inheritance and allelism of Russian wheat aphid resistance in several wheat lines. Plant Breeding 116:449-453 JOHNSON, J.J., AND QUICK, J.S. 1997. Colorado wheat performance test results. Colorado Wheat Farmer 39 (2):4-6. KHAN, Q.A., AND QUICK, J.S. 1997. Field and greenhouse evaluation of winter wheat for post anthesis drought stress using chemical desiccation. Agronomy Abstr., p. 76, Madison, WI MULAT, T.G., AND QUICK, J.S. 1997. Yield and yield components of hard white winter wheat lines in eastern Colorado. Agronomy Abstr., p. 72, ents emerge at CSU. Colorado Wheat Farmer 39 (1):8 QUICK, J.S. 1997. CSU releases two new Russian wheat aphid varieties. Colorado Wheat Farmer. 39 (3):4 QUICK, J.S., AND NISSEN, S. 1997. CSU developing herbicide resistant wheat seed. Colorado Wheat Farmer 39(3):1 ZHANG, Y., QUICK, J.S., AND S. LIU. 1997. Genetic variation in PI 294994 wheat for resistance to the Russian wheat aphid. Agronomy Abstr., p. 67, Madison, WI |
| 1998 |
DAVIS, J.G., AND QUICK, J.S. 1998. Nutrient management, and cultivar development and selection strategies to optimize water-use efficiency. J. Crop Prod. 1:221-240 DAVIS, J.G., AND QUICK, J.S. 1998. Nutrient management, and cultivar development and selection strategies to optimize water-use efficiency. pp. 221-240 In Nutrient Use in Crop production (Z . Rengel, ed.). The Haworth Press, Inc. New York ERKER, B. 1998. Utility of molecular marker technology and inheritance of resistance of PI 262660 in breeding Russian wheat aphid resistant wheats. M.S. Thesis. Colorado State University, Fort Collins, CO 53 pp IBRAHIM, A.M.H. 1998. Genetic analysis of heat tolerance in winter and spring wheat. Ph.D. Thesis. Colorado State University, Fort Collins, CO 80 pp KHAN, Q.A., AND QUICK, J.S. 1998. Chemical desiccation tolerance of winter wheat in the field and greenhouse. P. 134. Proc. 21st Hard Winter Wheat Workshop, Denver, CO LIU, S. 1998. Chromosome location of Russian wheat aphid resistance genes in wheat germplasms PI 372129, PI 243781, and PI 220127. M.S. Thesis. Colorado State University, Fort Collins, CO 36 pp MA, Z.Q., SAIDI, A., QUICK, J.S., AND LAPITAN, N.L.V. 1998. Genetic mapping of Russian wheat aphid resistance genes Dn2 and Dn4 in wheat. Genome 41:303-306 MULAT, T.G. 1998. Hard white winter wheat evaluation in eastern Colorado. M.S. Thesis. Colorado State University, Fort Collins, CO 110 pp MULAT, T.G., AND QUICK, J.S. 1998. Performance of hard white winter wheat lines in Colorado. p . 130. Proc. 21st Hard Winter Wheat Workshop, Denver, CO QUICK, J.S. 1998. CSU releases new `Prairie Red'. Colorado Wheat Farmer. 40 (2):3 QUICK, J.S. 1998. New developments in breeding for RWA resistance. p. 22. Proc. 21st Hard Winter Wheat Workshop, Denver, CO QUICK, J.S., AND JOHNSON, J.J. 1998. CSU variety trials aid planting decisions. Colorado Wheat Farmer 40(2):4-5. QUICK, J.S., AND PEAIRS, F.B. 1998. Breeding and germplasm development for Russian wheat aphid resistance. Proc. 9th Inter. Wheat Genetics Symposium, pp. 181-183. Saskatoon, Saskatchewan, Canada REYNOLDS, M.P., SINGH, R.P., IBRAHIM, A., AGEEB, O.A.A., LARQUE-SAAVEDRA, A., AND QUICK, J.S. 1998. Evaluating physiological traits to complement empirical selection in warm environments. Euphytica 100:85-94 SALMAN, A.A., AND QUICK, J.S. 1998. Desiccation tolerance and its association with assimilate partitioning in spring wheats for Colorado. Agron. Abstr.p.84 SALMAN, A.A., AND QUICK, J.S. 1998. Desiccation tolerance and its association with assimilate partitioning in spring wheats for Colorado. p. 132. Proc. 21st Hard Winter Wheat Workshop, Denver, CO |
| 1999 |
Clayshulte, S.R., Stromberger, J.A., Guira, A., Quick, J.S., Haley, S.D., Papa, D., and Zhang, Y. 1999. Production of doubled haploid plants through wheat x maize crosses and their integration in a breeding program. Agron. Abs. p. 80, Amer. Soc. of Agron., Madison, WI Haley, S.D., and Quick, J.S. 1998. Methodology for evaluation of chemical desiccation tolerance in winter wheat. Cer. Res. Commun. 26:73-79 Ibrahim, A.M., and Quick, J.S. 1999. Heritability of high temperature tolerance in winter and spring wheat. Agron. Abs. p. 71, Amer. Soc. of Agron., Madison, WI Liu, S., Zhang, Y., and Quick, J.S. 1999. Chromosome location of Russian wheat aphid resistance genes in wheat germplasms. Agron. Abs. p. 74, Amer. Soc. of Agron., Madison, WI |
| 2000 |
Haley, S.D. 2000. Enhanced wheat breeding program management with relational database software . P. 417. In Agron. Abstr. ASA, Madison, WI. Haley, S.D. 2000. Wheat breeding research fosters new developments. Colorado Wheat Farmer 42(2):7. Hesler, L.S., Riedell, W.E., Haley, S.D., Kieckhefer, R.W. and Collins, R.D. 2000. Resistance to Rhopalosiphum padi (L.) in wheat germplasm accessions. Proc. Intl. Plant Res. to Insects - 14th biennial workshop, Feb. 28-March 2, 2000, Fort Colliins, CO. Johnson, J., and Haley, S.D. 2000. Wheat decisions. Colorado Farmer-Stockman 54:39. Johnson, J., and Haley, S.D. 2000. Winter wheat variety performance trial results. Colorado Wheat Farmer 42(2):6. Mulat, T.G., Haley, S.D., and Quick, J.S. 2000. Chemical desiccation tolerance and stem reserve remobilization in hard red and hard white winter wheat lines. P. 105. In Agron. Abstr. ASA, Madison, WI. |
| 2001 |
Butler, J.D., Byrne, P.F., Nielsen, D.C., Doesken, N.J., McMaster, G.S., Haley, S.D., and Stushnoff, C. 2001. Determining temperature and precipitation variables that best explain Colorado wheat yields. In Agronomy abstracts. ASA, Madison, WI. Ibrahim, Amir H., and Quick, J. S. 2001. Genetic Control of High Temperature Tolerance in Wheat as Measured by membrane Thermal Stability. Crop Sci. 41:1405. Ibrahim, Amir H., and Quick, J. S. 2001. Heritability of Heat Tolerance in Winter and Spring Wheat. Crop Sci. 41:1401. Johnson, J., and Haley, S. D. 2001. Winter wheat performance. Colorado Farmer-Stockman 55(11):19-23. Khan, Q. A., and Quick, J. S. 2001. Cell Membrane Thermostability and its Relation to Grain Yield in Spring Wheat. In Agronomy Abstracts. ASA, Madison, Wisconsin. McMaster, G.S., Byrne, P.F., Haley, S.D., Nielsen, D.C., Stushnoff, C., and Doesken, N.J. 2001 . Winter wheat varieties differ in phenological and canopy development responses to water stress. In Agronomy abstracts. ASA, Madison, WI. Quick, J.S., Haley, S.D., Stromberger, J.A., Clayshulte, S., Clifford, B., Johnson, J.J., Peairs, F.B., Rudolph, J.B., and Lorenz, K. 2001. Registration of 'Prowers 99' wheat. Crop Sci . 41:929. |
| 2002 |
Ball, E.L. 2002. Inheritance and chromosomal location of wheat streak mosaic virus resistance in wheat. M.S. Thesis, Colorado State University, Fort Collins, CO. Bariana, H.S., Brown, G.N., Ahmed, N.U., Khatkar, S., Conner, R.L., Wellings, C.R., Haley, S.D ., Sharp, P.J., and Laroche, A. 2002. Characterisation of Triticum vavilovii-derived stripe rust resistance using genetic, cytogenetic and molecular analyses and its marker-assisted selection. Theor. Appl. Genet. 104:315-320. Berzonsky, W.A., Ding, H., Haley, S.D., Harris, M.O., Lamb, R.J., McKenzie, R.I.H., Ohm, H.W., Patterson, F.L., Peairs, F.B., Porter, D.R., Raticliffe, R.H., and Shanower, T.G. 2002. Breeding wheat for resistance to insects. p. 221-296 In J. Janick (ed.) Plant Breeding Reviews , John Wiley & Sons, Hobeken, NJ. Haley, S.D., Martin, T.J., Quick, J.S., Seifers, D.L., Stromberger, J.A., Clayshulte, S.R., Clifford, B.L., Peairs, F.B., Rudolph, J.B., Johnson, J.J., Gill, B.S., and Friebe, B. 2002. Registration of CO960293-2 wheat germplasm resistant to wheat streak mosaic virus and Russian wheat aphid. Crop Sci. 42:1381-1382. Johnson, J., Haley, S.D., and Westra, P. 2002. The CLEARFIELD* production system for improved weed control in Colorado winter wheat - a fact sheet for Colorado wheat producers. CSU Coop. Ext. Service Fact Sheet No. 3.116. Thomas, J., Hein, G., Baltensperger, D., Nelson, L., and Haley, S.D. 2002. Managing the Russian wheat aphid with resistant wheat varieties. Univ. Nebraska Fact Sheet NF 96-307 (revised September 2002). Yu, L., Haley, S.D., Perret, J., Harris, M., and Qian, M. 2002. Antioxidant properties of hard winter wheat extracts. Food Chem. 78:457-461. Yu, L., Haley, S.D., Perret, J., Harris, M., Wilson, J., and Qian, M. 2002. Free radical scavenging properties of wheat extracts. J. Agric. Food Chem. 50:1619-1624. |
| 2003 |
Haley, S.D., Lazar, M.D., Quick, J.S., Johnson, J.J., Peterson, G.L., Stromberger, J.A., Clayshulte, S.R., Clifford, B.L., Pester, T.A., Nissen, S.J., Westra, P.H., Peairs, F.B., and Rudolph, J.B. 2003. Above winter wheat. Can. J. Plant Sci. 83:107-108. Haley, S.D., Peairs, F.B., Rudolph, J.B., and Walker, C.B. 2003. Occurrence of a new Russian wheat aphid biotype in Colorado. Proc. West. Crop Sci. Soc. of Amer. Ann. Meetings, 23-24 Jun. 2003, Fort Collins, CO. Haley, S.D., Quick, J.S., Martin, T.J., Johnson, J.J., Peairs, F.B., Stromberger, J.A., Clayshulte, S.R., Clifford, B.L., and Rudolph, J.B. 2003. Registration of 'Avalanche' wheat. Crop Sci. 43:432. Hesler, L.S., Riedell, W.E., Kieckhefer, R.W., and Haley, S.D. 2003. Responses of Rhopalosiphum padi on cereal aphid-resistant wheat accessions. J. Agric. Urban Entomol. 19:133-140. Johnson J.J., Berrada, A., Bosley, B., Bruns, R., Davis, J., Dillon, M., Haley, S.D., Hain, J. , Hanavan, D., Hill, J., Johnson, C.L., Khosla, R., Larsen, K., Peairs, F., Pearson, C., Schweissing, F., Stack, M., Sumpter, C., Thompson, C., Thompson, D., Westfall, D., and Westra, P. 2003. Making Better Decisions, 2002 Colorado Winter Wheat Performance Trials. Colorado State University Agric. Exp. Stn. Technical Report TR 03-4. Johnson J.J., Berrada, A., Dillon, M., Haley, S.D., Pearson, C.H., Quick, J.S., and Stack, M. 2003. Making Better Decisions, 2002 Colorado Spring Wheat, Barley, and Oats Variety Performance Trials. TR 03-2. Lapitan, N.L.V., Peng, J.P., Tahir, M., Sharma, V., and Haley, S.D. 2003. High throughput marker assisted selection in wheat: breeding for Russian wheat aphid resistance. Proc. 10th Intl. Wheat Gen. Symp. 2:745-747. Lazar, M.D., Haley, S.D., Quick, J.S., Johnson, J.J., Peterson, G.L., Stromberger, J.A., Clayshulte, S.R., Clifford, B.L., Pester, T.A., Nissen, S.J., Westra, P.H., Peairs, F.B., and Rudolph, J.B. 2003. AP502 CL winter wheat. Can. J. Plant Sci. 83:109-110. McMaster, G.S., Ascough, J.C., Shaffer, M.J., Deer-Ascough, L.A., Byrne, P.F., Nielsen, D.C., Haley, S.D., Andales, A.A., and Dunn, G.H. 2003. GPFARM plant model parameters: complications of varieties and the genotype x environment interaction in wheat. Trans. Amer. Soc. Agri. Eng. (ASAE) 46:1337-1346. Pardina-Malbran, F., Johnson, J.J., Cipra, J., Haley, S.D., and Reich, R. 2003. Predicting wheat yield and quality traits using spatial and environmental models. In Agronomy abstracts. ASA, Madison, WI. Randolph, T.L., Rudolph, J.B., Peairs, F.B., Haley, S.D., and Walker, C.B. 2003. Occurrence of a new biotype of Russian Wheat Aphid in Colorado. Proc. Entom. Soc. of Amer. Ann. Meetings, 26-29 Oct. 2003, Cincinnati, OH. Yu, L., Perret, J., Harris, M., Wilson, J., and Haley, S.D. 2003. Antioxidant properties of bran extracts from Akron wheat grown at different locations. J. Agric. Food Chem. 51:1566-1570 . |
| 2004 |
Butler, J.D., Byrne, P.F., Chapman, P.L., and Haley, S.D. 2004. Agronomic performance of Rht alleles in a spring wheat population across a range of moisture levels. Proc. National Wheat Workers Workshop, Kansas City, MO, 23-24 Feb. 2004. Clayshulte, S.R., Haley, S.D., Chapman, P.L., Pardina-Malbran, F., Seabourn, B.W., and Chung, O.K. 2004. Clustering of nursery locations based on hard winter wheat regional quality evaluations. Proc. National Wheat Workers Workshop, Kansas City, MO, 23-24 Feb. 2004. Collins, M.B. 2004. Reaction of spring wheat genotypes and wheat germplasm accessions to Russian wheat aphid biotypes 1 and 2. M.S. Thesis, Colorado State University, Fort Collins, CO . 69 pp. Collins, M.B., Haley, S.D., Randolph, T.L., Peairs, F.B., and Chapman, P.L. 2004. Field Reaction of Dn4- and Dn7-carrying spring wheat genotypes under artificial Russian wheat aphid biotype 1 infestation. In Agronomy abstracts. ASA, Madison, WI. Collins, M.B., Haley, S.D., Rudolph, J., Randolph, T., and Peairs, F.B. 2004. Russian wheat aphid biotype B resistance among wheat germplasm accessions. Proc. National Wheat Workers Workshop, Kansas City, MO, 23-24 Feb. 2004. Haley, S.D., Peairs, F.B., Walker, C.B., Rudolph, J.B., and Randolph, T.L. 2004. Occurrence of a new Russian wheat aphid biotype in Colorado. Crop Sci. 44:1589-1592. Haley, S.D., Quick, J.S., Johnson, J.J., Peairs, F.B., Stromberger, J.A., Clayshulte, S.R., Clifford, B.L., Rudolph, J.B., Chung, O.K., and Seabourn, B.W. 2004. Registration of 'Ankor' wheat. Crop Sci. 44:1025-1026. Johnson J.J., and Haley, S.D. 2004. 2004 Colorado winter wheat variety performance trial results. Agronomy News, From the Ground Up Extension Newsletter, Vol.24, Issue 2. Soil and Crop Sciences Department, Colorado State University. Johnson J.J., and Haley, S.D. 2004. A decision guide for winter wheat variety selection-risk reduction through variety maturity ratings. Agronomy News, From the Ground Up Extension Newsletter, Vol. 24, Issue 2. Soil and Crop Sciences Department, Colorado State University. Johnson, J.J., Blunt, T., Bruns, R., Fichtner, S., Haley, S.D., Hanavan, D., Hill, J.P., Johnson, C., Peairs, F., Pearson, C., Pollart, M., Schwartz, H.F., Sears, R., Velasco, V., and Westra, P. 2004. Making Better Decisions, 2003 Colorado Winter Wheat Variety Performance Trials. Colorado State University Agric. Exp. Stn. Technical Report TR 04-3. Mulat, T.G. 2004. Chemical desiccation tolerance and nonstructureal carbohydrate dymamics in winter wheat. Ph.D. Dissertation, Colorado State University, Fort Collins, CO. 111 pp. Pardina-Malbran, F. 2004. Spatial and environmental modeling of wheat yield and quality in Colorado using on-farm trials and GIS. M.S. Thesis, Colorado State University, Fort Collins, CO. 97 pp. Tahir, M., Sharma, V., Haley, S.D., Butler, J., and Lapitan, N.L.V. 2004. High density mapping of a rye gene (Dn7) conferring resistance to Russian wheat aphid in wheat. Proc. Plant and Animal Genome, San Diego, CA. Yu, L., Haley, S.D., Perret, J., and Harris, M. 2004. Comparison of wheat flours grown at different locations for their antioxidant properties. Food Chem. 86:11-16. |
| 2005 |
Butler, J.D., Byrne, P.F., Mohammadi, V., Chapman, P.L., and Haley, S.D. 2005. Agronomic performance of Rht alleles in a spring wheat population across a range of moisture levels. Crop Sci. 45:939-947. Butler, J.D., Haley, S.D., and Seabourn, B.W. 2005. Efficiency of selection for wheat kernel characteristics using near-infrared reflectance spectroscopy. In Agronomy abstracts. ASA, Madison, WI. Clayshulte, S.R., Haley, S.D., Byrne, P.F., Dabbert, T., and Shan, X. 2005. Agronomic trait associations and allelic diversity at the Xgwm261 marker locus in winter wheat. In Agronomy abstracts. ASA, Madison, WI. Collins, M.B., Haley, S.D., Peairs, F.B., and Rudolph, J.B. 2005. Russian wheat aphid Biotype 2 resistance among wheat germplasm accessions. Crop Sci. 45:1877-1880. Collins, M.B., Haley, S.D., Randolph, T.L, Peairs, F.B., and Rudolph, J.B. 2005. Comparison of Dn4- and Dn7-carrying spring wheat genotypes artificially infested with Russian wheat aphid (Homoptera: Aphididae) Biotype 1. J. Econ. Entomol. 98(5):1698-1703. Haley, S.D., Quick, J.S., Johnson, J.J., Peairs, F.B., Stromberger, J.A., Clayshulte, S.R., Clifford, B.L., Rudolph, J.B., Seabourn, B.W., Chung, O.K., Jin, Y., and Kolmer, J. 2005. Registration of 'Hatcher' wheat. Crop Sci. 45:2654-2655. Johnson J.J., and Haley, S.D. 2005. 2005 Colorado winter wheat variety performance trial results. Agronomy News, From the Ground Up Extension Newsletter, Vol. 25, Issue 1. Soil and Crop Sciences Department, Colorado State University. unpublished Johnson J.J., and Haley, S.D. 2005. Winter wheat variety selection in Colorado for Fall 2005. Agronomy News, From the Ground Up Extension Newsletter, Vol. 25, Issue 1. Soil and Crop Sciences Department, Colorado State University. unpublished Randolph, T.L., Peairs, F.B., Koch, M., Walker, C.B., Stubbs, J.R., Quick, J.S., and Haley, S .D. 2005. Yield response and categories of resistance to Russian Wheat Aphid in four Dn4 hard red winter wheats. J. Econ. Entomol. 98:588-594. Stromberger, J.A., Stushnoff, C., Butler, J.D., Clifford, B.L., and Haley, S.D. 2005. Genotype and environment effects and quality trait associations with antioxidant properties of wheat grain. In Agronomy abstracts. ASA, Madison, WI. |
| 2006 |
Burd, J.D., Porter, D.R., Puterka, G.J., Haley, S.D., and Peairs. F.B.. 2006. Biotypic variation among North American Russian wheat aphid (Homoptera: Aphididae) populations. J. Econ . Entomol. 99:1862-1866. Haley, S.D. 2006. Winter wheat breeding and genetics program. 2004 Southwest Colorado Research Center Research Report. Colorado State University Agric. Exp. Stn. Technical Report TR 06-3. Haley, S.D., and Johnson, J.J. 2006. Colorado wheat variety database now available for download. Colorado Wheat Farmer. Vol. 48 (2):7. Haley, S.D., DeLacy, I.H., and Johnson, J.J.. 2006. Multilocational (1990-2006) analysis of Colorado winter wheat yield trial data. In Agronomy abstracts. ASA, Madison, WI. Haley, S.D., Johnson, J.J., Peairs, F.B., Quick, J.S., Westra, P.H., Stromberger, J.A., Clayshulte, S.R., Clifford, B.L., Rudolph, J.B., Giura, A., Seabourn, B.W., Chung, O.K., Jin, Y., and Kolmer, J.. 2006. Registration of 'Protection' wheat. Crop Sci. 46:995-996. Haley, S.D., Johnson, J.J., Peairs, F.B., Quick, J.S., Westra, P.H., Stromberger, J.A., Clayshulte, S.R., Clifford, B.L., Rudolph, J.B., Giura, A., Seabourn, B.W., Chung, O.K., Jin, Y., and Kolmer, J.. 2006. Registration of 'Bond CL' wheat. Crop Sci. 46:993-994. Ibrahim, A.M.H., Haley, S.D., Jin, Y., Langham, M.A.C., Rickertsen, J., Kalsbeck, S., Little, R., Chung, O.K., Seabourn, B.W., and McVey, D.V.. 2006. Registration of 'Wendy' Wheat. Crop Sci. 36:1389-1390. Johnson, J.J., and Haley, S.D. 2006. Making better decisions - Winter wheat variety selection in Colorado for Fall 2006. Colorado Wheat Farmer. Vol. 48 (2):7. Johnson, J.J., and Haley, S.D. 2006. Variety selection. p. 9-15. In F.B. Peairs (ed.) Colorado wheat production handbook. Colorado Association of Wheat Growers, Denver, CO. Johnson, J.J., Haley, S.D., Hanavan, D., Peairs, F., Stack, M., Tisserat, N., Berrada, A., Walker, T., Randolph, T., Rudolph, J., Hammon, B. 2006. Making Better Decisions, 2005 Colorado Winter Wheat Variety Performance Trials. Colorado State University Agric. Exp. Stn. Technical Report TR 06-09. Puterka, G.J., Burd, J.D., Mornhinweg, D.W., Haley, S.D., and Peairs, F.B.. 2006. Response of resistant and susceptible barley to infestations of five Diuraphis noxia (Homoptera: Aphididae) biotypes. J. Econ. Entomol. 99: 2151-2155. Seifers, D.L., Martin, T.J., Harvey, T.L., Haber, S., and Haley, S.D.. 2006. Temperature sensitivity and efficacy of Wheat streak mosaic virus resistance derived from CO960293 wheat. Plant Dis. 90:623-628. Shan, X., Clayshulte, S.R., Byrne, P.F., and Haley, S.D.. 2006. Variation for glutenin and waxy alleles in U.S. hard winter wheat germplasm. In Agronomy abstracts. ASA, Madison, WI. Zheng, S. X. Shan, P.F. Byrne, and S.D. Haley. 2006. Effects of glutenin subunit variation and 1RS.1AS translocation on dough properties of wheat grown in Colorado environments. In Agronomy abstracts. ASA, Madison, WI. |